Ecology, Evolution, and Marine Biology (EEMB)

EEMB 1122. Physical Oceanography. 4 Hours.

Provides a description of the physical properties and composition of seawater, waves, tides, and ocean currents. Discusses how these properties are measured by oceanographers and how they influence the earth’s environment and climate.

EEMB 1123. Biological Oceanography. 4 Hours.

Covers the productivity of plant and animal life in the various zones of the ocean and the growing economic importance of the oceans as a source of food for the expanding world population.

EEMB 1145. Beginning Scuba. 1 Hour.

Focuses on basic skin diving and scuba diving skills, with emphasis on safety. Requires lab fee. Requires ability to pass a swim test and basic comfort in the water.

EEMB 1450. Introduction to Marine Biology. 4 Hours.

Surveys the tremendous diversity of marine organisms in the context of the major marine ecosystems in which they are found. Explores interactions among organisms and how the physical and chemical environment influence marine organisms. Links changes on land to declines in organism numbers and diversity and explores the benefits humans gain from our relationship with the marine environment. Offers opportunities to investigate recent advances and understanding of marine organisms and their environments.

EEMB 1990. Elective. 1-4 Hours.

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.

EEMB 2290. Ecology and Evolution of Behavior. 4 Hours.

Studies fundamental biological principles at behavioral, ecological, and evolutionary levels. Covers ethology, ecology, genetics, and comparative psychology, all within the conceptual framework of evolutionary theory. Explores both scientific practice and progress through readings, discussion, and projects. Illustrates the process by which biologists study questions about the evolutionary origin of behavior through a series of in-class activities, computer modeling assignments, interpretation of graphical data, collection and statistical analyses of behavioral data, as well as the generation and presentation of research. Does not focus on the neurological basis of behavior. Offers students an opportunity to become critical thinkers, critical readers, and to attain tools to interpret the world in a unique way. Requires permission of advisor.

EEMB 2302. Ecology. 4 Hours.

Offers students an opportunity to learn about the environmental and biological processes that control the distribution and abundance of species and controlling factors that operate on individuals, populations, and communities. The lecture and laboratory introduce a set of generalizable concepts that are of fundamental importance to plant and animal life on the land and in the sea and provide hands-on experiential learning that reinforce concepts covered in lecture. Offers students an opportunity to become proficient in the following: (a) understanding research results the primary literature; (b) conducting a research experiment; (c) interpreting the results of in-class research; (d) communicating results as manuscript.

EEMB 2303. Lab for EEMB 2302. 1 Hour.

Accompanies EEMB 2302. Covers topics from the course through various experiments.

EEMB 2400. Introduction to Evolution. 4 Hours.

Introduces evolutionary thinking, including contemporary examples of evolution. To understand the evolution of Charles Darwin’s “endless forms most beautiful,” the course adopts an integrative approach that includes information from ecology, genetics, molecular biology, biogeography, and paleobiology. Considers mechanisms of evolutionary change—how does it happen? Examines adaptation, the process by which attributes of an organism change to enhance fitness and the evolutionary history of life on our planet—what was the first living thing, how does speciation occur, what have we learned about evolution of life in the distant past, and how did humans evolve. Includes student presentations and analysis of scientific literature.

EEMB 2410. Fish Biology and Ecology. 4 Hours.

Covers fish evolutionary relationships, functional morphology, global biogeography, reproductive behavior, and basic ecology. Considers how fishes interact with each other and with their environment across multiple scales. Focuses on how basic life requirements such as habitat use, behavior, foraging, and reproduction lead to variation among individuals, affect population dynamics, and impact the structure and function of community organization and ultimately how these processes influence broad-scale patterns and dynamics at the ecosystem level.

EEMB 2411. Lab for EEMB 2410. 1 Hour.

Accompanies EEMB 2410. Covers topics from the course through various experiments.

EEMB 2420. Fisheries Biology, Policy, and Conservation. 4 Hours.

Focuses on the study and management of economically valuable fish species. Studies the basic biology and ecology of fisheries species, quantifying and modeling their population biology to their interactions with each other and the environment. Requires students to read and analyze the scientific literature, to complete worksheets and writing assignments, and to develop and present research projects. Covers traditional stock assessment methods as well as how fisheries science and management has evolved more recently to integrate community- and ecosystem-level information. Reviews fisheries and how fishers are managed, their involvement in the management process, and the future fisheries in the United States and elsewhere.

EEMB 2610. Plant Biology. 4 Hours.

Examines the biology and diversity of plants and plant-like organisms. Explores the relationships between humans and plants by looking at plants through three different perspectives: feeding a starving world; curing a sick world; and engineering a better world. Employs case studies to highlight major themes.

EEMB 2611. Lab for EEMB 2610. 1 Hour.

Accompanies EEMB 2610. Covers topics from the course through various experiments.

EEMB 2616. Invertebrate Zoology. 4 Hours.

Surveys the tremendous diversity of invertebrates, emphasizing their form and function in ecological and evolutionary contexts. Explores functional morphology, systematics, phylogenetic relationships, ecology, and economic importance of the major invertebrate phyla. Discusses comparisons among phyla to enhance understanding of evolutionary relationships.

EEMB 2617. Lab for EEMB 2616. 1 Hour.

Accompanies EEMB 2616. Covers topics from the course through various experiments.

EEMB 2618. Vertebrate Zoology. 4 Hours.

Explores functional morphology, systematics, ecology, and phylogenetic relationships of the major vertebrate phyla.

EEMB 2619. Lab for EEMB 2618. 1 Hour.

Accompanies EEMB 2618. Covers topics from the course through various experiments.

EEMB 2700. Marine Biology. 4 Hours.

Examines biological aspects of natural ocean ecosystems and the physical processes that regulate them. Covers distributions, abundances, and interactions of marine organisms; interactions between organisms and the transformation and flux of energy and matter in marine ecosystems; and aspects of physiology related to marine species distributions, abundances, and roles. Students generate, evaluate, discuss, and present data from primary research and apply their knowledge of the scientific method and biological concepts through the creation of a written grant proposal.

EEMB 2701. Lab for EEMB 2700. 1 Hour.

Accompanies EEMB 2700. Covers topics from the lecture course through discussions and experiments.

EEMB 2990. Elective. 1-4 Hours.

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.

EEMB 3120. Physical Biology of Marine Organisms. 4 Hours.

Introduces principles from the physical sciences (fluid and solid mechanics, mass and heat transfer theory) applied to the analysis of form, function, ecology, and evolution of marine organisms. Topics covered include suspension and deposit feeding in invertebrates, allometry of metabolic processes, drag and lift in sessile organisms, locomotion of nekton (fishes, marine mammals) and plankton, diffusive limitations to metabolic transactions in marine invertebrates and algae, thermal transactions in intertidal organisms, the biology of the benthic boundary layer, and the properties of biomaterials and biological structures. Presents engineering methods and measurement techniques applicable to biomechanical investigations.

EEMB 3450. Physiological Adaptations to the Environment. 4 Hours.

Explores the evolutionary mechanisms by which organisms adapt physiologically to survive, and thrive, in diverse, often seemingly “hostile,” habitats. Examines paleo- and modern examples of adaptation with the goal of predicting species success or failure as our planetary environment changes rapidly. Topics include adaptation of cellular metabolism, adaptations to variable oxygen availability and to changes in pH, the roles of water and microsolutes in regulation of the internal environment of cells, and the effects of temperature on cellular function and the biogeographic distribution of organisms. Includes student presentations and analysis of scientific literature. Requires junior or senior standing; sophomores admitted by permission of instructor; EEMB 2400 or ENVR 2400 recommended but not required.

EEMB 3455. Ecosystems Ecology. 4 Hours.

Focuses on the foundational principles of ecosystems ecology. Examines the flow of energy and materials through both the biosphere (plants, animals, and microbes) and the geosphere (soils, atmospheres, and oceans) and the role that humans are playing in altering these key fluxes. Studies elemental cycles that are critically important for human and environmental sustainability—including carbon, nitrogen, and phosphorus—and examines similarities and differences in these cycles and flows while drawing on examples from both terrestrial and marine systems. Seeks to understand how changes in ecosystem structure ultimately affect ecosystem function and how this translates into the important services ecosystems provide.

EEMB 3460. Conservation Biology. 4 Hours.

Explores conservation biology, an interdisciplinary science that focuses on conservation of biological diversity at multiple levels. Emphasizes the causes and consequences of biodiversity loss and demonstrates how ecological and evolutionary principles are applied to conservation problems. Covers sustainability; climate change; introduced species; conservation of threatened and endangered species; and pollution, disease, and habitat restoration using examples from marine, aquatic, and terrestrial systems. Offers students an opportunity to read, discuss, evaluate, and present data from primary research through written assignments and oral debates and to apply this knowledge to conservation issues. Emphasizes critical thinking, problem solving, and recognizing multiple perspectives.

EEMB 3465. Ecological and Conservation Genetics. 4 Hours.

Offers an overview of ecological and conservation genetics, an interdisciplinary science that focuses on understanding the processes that determine genetic diversity at the individual to population level. Focuses on fundamental concepts in evolutionary ecology and population and quantitative genetics, then applies those concepts to solving real-world problems in conservation science. Covers harvested populations, inbreeding, climate change, introduced species, conservation of threatened and endangered species, adaptation, and habitat restoration. Exposes students to multiple sides of these issues and the science that underpins them. Offers students an opportunity to develop the R programming skills required to analyze the complex data sets that often emerge when addressing cutting-edge questions in genetics. Includes writing and coding exercises and mathematical derivations. Emphasizes critical thinking and problem solving.

EEMB 3475. Terrestrial Wildlife Ecology. 4 Hours.

Discusses wildlife ecology and management, mainly focusing on terrestrial species. Topics include habitat use, behavior, wildlife conservation, parasites and pathogens, wildlife sampling, and wildlife management. Offers students an opportunity to participate in activities in which they look at and interpret wildlife data. Course format includes group work, analyzing the scientific literature, and in-class activities. Requires sophomore or junior standing; open to seniors with permission of instructor.

EEMB 3990. Elective. 1-4 Hours.

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.

EEMB 4001. Landscape and Restoration Ecology. 4 Hours.

Topics include ecosystem processes, spatial patterns, disturbance, species distributions, invasive species, and habitat loss. Offers students an opportunity to participate in activities in which they look at and interpret spatial data. Course format includes group work, analyzing the scientific literature, and in-class activities. Requires sophomore or junior standing; open to seniors with permission of instructor.

EEMB 4010. Mammalogy. 4 Hours.

Surveys the mammals of the world, including their evolution, morphology, physiology, behavior, and ecology. Students conduct a research project in which they investigate the morphology, evolution, ecology, and behavior of a species and present their findings to the class. Includes reading and analyzing the scientific literature and conducting in-class activities.

EEMB 4990. Elective. 1-4 Hours.

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.

EEMB 4992. Directed Study. 1-4 Hours.

Offers independent work under the direction of members of the department on a chosen topic. Course content depends on instructor. May be repeated without limit.

EEMB 5130. Ecological Dynamics. 4 Hours.

Offers a comprehensive overview of mathematical and computational concepts needed to construct (meta)population, (meta)community, and (meta)ecosystem models. Focuses on how to mathematically derive and model processes (growth, trophic and nontrophic species interactions, dispersal, and environmental variability) to understand patterns of population abundance and species diversity. Emphasizes the mathematical tools required to analyze the dynamical behavior of ecological models (stability, invasion, graphical, and numerical analyses) and validate model predictions using empirical data (via maximum likelihood and optimization methods). Sophomores admitted by permission of instructor.

EEMB 5131. Lab for EEMB 5130. 1 Hour.

Accompanies EEMB 5130. Offers supervised lab sessions designed to show how the topics covered in the lectures can be addressed in industry-standard programming environments.

EEMB 5303. Marine Biology Careers Seminar. 1 Hour.

Covers the information and tools needed to begin pursuing career opportunities in marine biology. Encourages students to explore a variety of career paths, construct résumés, contact potential employers for their internship and permanent positions. Presents invited speakers from state and federal agencies, and from private consulting firms, to talk about their work and career track.

EEMB 5504. Biology of Corals. 3 Hours.

Focuses on the biology of Scleractinian reef-building corals and associated anthozoans found in coral reef ecosystems. Topics include systematics, anatomy, physiology, and population biology of corals, with an emphasis on the latest techniques employed by coral molecular biologists and physiologists.

EEMB 5506. Biology and Ecology of Fishes. 3 Hours.

Presents an examination of the systematics, functional morphology, and behavioral, larval, and community ecology of reef fishes through lectures. Field and laboratory experiments focus on morphology, behavior, and community ecology of reef fishes.

EEMB 5508. Marine Birds and Mammals. 2 Hours.

Studies principles of classification, anatomy, physiology, behavior, and evolution of seabirds and marine mammals. Also addresses conservation and protection of animals and essential habitat. Includes field trips to observe local species.

EEMB 5509. Lab for EEMB 5508. 1 Hour.

Accompanies EEMB 5508. Covers topics from the course through various experiments.

EEMB 5511. Adaptations of Aquatic Organisms. 3 Hours.

Explores the adaptive responses of marine organisms to variations in environmental factors. Focuses on physiological responses to a variety of natural and anthropogenic conditions. The laboratory component includes a combination of field and laboratory experiments.

EEMB 5512. Tropical Terrestrial Ecology. 1 Hour.

Studies the animals, plants, and ecosystems of the new world tropics, with the community structure and diversity of terrestrial Jamaican habitats as an example. Includes field trips to lowland forests, carbonate caves, and the Blue Mountain mist-montane forest. The issue of land use and development vs. conservation is a recurring theme.

EEMB 5514. Marine Ecology. 4 Hours.

Examines processes and interactions in ocean ecosystems. Topics include an introduction to major ocean ecosystems; the biotic and abiotic factors influencing the distributions, abundances, and interactions of marine organisms; and the transformation and flux of energy and matter in marine systems. Particularly emphasizes local coastal habitats, which are used to demonstrate quantitative field research methods.

EEMB 5515. Lab for EEMB 5514. 1 Hour.

Accompanies EEMB 5514. Covers topics from the course through various experiments.

EEMB 5516. Oceanography. 4 Hours.

Offers an integrated overview of physical, chemical, biological, and geological processes operating in the world ocean. Seemingly unrelated topics like plate tectonics, oscillating currents and waves in the atmosphere, the activities of microbes and phytoplankton, and land-use practices in the middle of the continent have global reach and interact with each other in surprising yet understandable ways. Examines how new technologies have allowed stunning insights into global weather and climate, the deep sea, biodiversity, and how the biogeochemistry of the oceans can be measured and understood. Presents data use and analysis and formal reasoning used in marine science. Views the ocean as a “system of systems” where integration of experience from disparate disciplines is key.

EEMB 5517. Lab for EEMB 5516. 1 Hour.

Accompanies EEMB 5516. Offers experiential field and laboratory exercises in oceanography. The New England rocky intertidal, subtidal, wetlands, barrier islands, and dunes provide opportunities for field exercises in marine geology, physical oceanography, and marine ecology. Investigates processes affecting changes in the global ocean, such as ocean acidification; temperature stress in organisms; hydrodynamic drag and lift; suspension feeding; and the ecophysiology of reef corals, boreal invertebrates, and macroalgae.

EEMB 5518. Ocean and Coastal Processes. 2 Hours.

Examines the coupling between physical and biological processes on coral reefs and adjacent habitats. Focuses on biophysical, oceanographic, and benthic-pelagic processes acting in coral reef and associated nearshore ecosystems. Specific topics include oceanographic forcing mechanisms, organismal biomechanics, hydrodynamics, and nutrient dynamics.

EEMB 5520. Coral Reef Ecology. 2 Hours.

Examines the ecology and paleoecology of coral reefs. This course highlights the ecological importance of coral reefs and associated nearshore communities, ecosystem function, changes in reef biotas through geologic time, and the causes and consequences of reef degradation worldwide.

EEMB 5522. Experimental Design Marine Ecology. 4 Hours.

Includes introduction to and application of observational methods in three local marine habitats, experimental design, statistical analysis, R statistical computing and graphics software, and principles of marine ecology. Combines lecture, hand-on research experience, and computer laboratory and includes reading and analyzing the scientific literature and developing research projects. At the end of the semester, students are expected to demonstrate an integrative mastery of course topics by writing a scientific manuscript about a class experiment. Seeks to prepare students for practicing ecology in new environments and to provide students with the foundational knowledge necessary for pursuing more complex concepts in experimental design, statistical analysis, and marine ecology.

EEMB 5523. Lab for EEMB 5522. 1 Hour.

Accompanies EEMB 5522. Covers topics from the course through various experiments.

EEMB 5524. Molecular Marine Biology. 3 Hours.

Uses molecular approaches (electrophoresis and DNA) to determine genetic relationships at the population and species level for the study of ecological and evolutionary questions. Techniques learned are applied to research projects.

EEMB 5526. Marine Microbial Ecology. 2 Hours.

Examines the diversity of marine microorganisms and recent advances in the area of microbial ecology. Emphasizes the structure and function of microbial food webs in marine communities.

EEMB 5527. Lab for EEMB 5526. 1 Hour.

Accompanies EEMB 5526. Covers topics from the course through various experiments.

EEMB 5528. Marine Conservation Biology. 3 Hours.

Examines several critical issues facing marine ecosystems, including invasive species, marine pollution and eutrophication, fisheries impacts, physical alteration of habitats, and global climate change. Offers students an opportunity to spend field time surveying intertidal and subtidal habitats within the San Juan Islands and Friday Harbor Marine Reserve and to conduct independent research projects.

EEMB 5530. Molecular Ecology and Evolution. 4 Hours.

Exposes students to the molecular techniques and analyses used to examine the genetic relationships among individuals, populations, and species.

EEMB 5531. Lab for EEMB 5530. 1 Hour.

Accompanies EEMB 5530. Covers topics from the course through various experiments.

EEMB 5532. Physiological and Molecular Marine Ecology. 3 Hours.

Explores the physiological responses of marine organisms to variations in environmental factors. Uses complementary techniques, including molecular and physiological approaches, to determine genetic relationships at the species and population level and elucidate the mechanistic basis of organismic responses to environmental conditions at the level of genes and gene products.

EEMB 5534. Marine Invertebrate Zoology and Botany. 4 Hours.

Surveys the major groups of marine invertebrates, algae, and plants, in addition to their ecological roles and relationships. Offers students an opportunity to learn to identify these groups and understand the mechanisms they use to survive and adapt to changing oceans. Topics include ecological and evolutionary importance, ecosystem engineering, adaptive physiology, and climate change effects. Emphasizes interrelationships among major taxa. Hands-on learning includes field identification; visits to intertidal and subtidal marine environments; and specimen dissection, preparation, and cataloging. Offers students an opportunity to improve skills in reading and discussing scientific literature, experimental design, and scientific communication. Restricted toThree Seas students only; not open to students who have taken EEMB 5500 or EEMB 5502.

EEMB 5535. Lab for EEMB 5534. 1 Hour.

Accompanies EEMB 5534. Covers topics from the course through various experiments.

EEMB 5536. Ocean and Coastal Sustainability. 3 Hours.

Offers students advanced training in the expanding field of sustainability, with a combined focus on the practical aspects of systems management and the theoretical understanding of whole-systems design and resiliency. Seeks to train future leaders capable of creating innovative solutions to sustainability issues at local and global levels. Key interdisciplinary themes discussed include the social and political aspects of ocean and coastal sustainability (i.e., education and communication), sustainable development and ecosystem stability, the impacts of climate change on ocean and coastal resilience, and the economic and entrepreneurial possibilities in the field of sustainability. Restricted to Three Seas students only.

EEMB 5548. Sociobiology. 4 Hours.

Studies sociobiology, a field of biology that strives to understand the biological basis of social behavior in animals. Sociobiology is a multidisciplinary science, meshing together ethology (animal behavior), ecology, genetics, population biology, and comparative psychology, all within the conceptual framework of evolutionary theory. Why do animals live in societies? Why do animals cooperate? Why do they sometimes show extreme forms of altruism? What are the costs and benefits of group living? Reviews studies on nonhuman animals that demonstrate sociobiological principles by using a series of in-class activities, computer modeling assignments, interpretation of graphical and tabulated data, collection and statistical analyses of behavioral data, as well as the generation and presentation of research.

EEMB 5560. Entomology. 4 Hours.

Studies the biology of insects and related arthropods including their anatomy, morphology, physiology, development, taxonomy, ecology, behavior, and life histories. Includes field and laboratory study of insect biology.

EEMB 5561. Lab for EEMB 5560. 1 Hour.

Accompanies EEMB 5560. Covers topics from the course through field and laboratory study, including insect collection.

EEMB 5562. Herpetology. 4 Hours.

Offers a survey of the amphibians and reptiles of the world, with emphasis on eastern North America. Topics include morphology, physiology, systematics, paleontology, ecology, zoogeography, and behavior. Includes field trips to observe the habitats and behavior of local herpetofauna. Laboratory emphasizes systematics and ecology.

EEMB 5563. Lab for EEMB 5562. 1 Hour.

Accompanies EEMB 5562. Covers topics from the course through various experiments.

EEMB 5564. Ornithology. 4 Hours.

Offers a survey of the birds of the world including morphology, physiology, systematics, behavior, ecology, zoogeography, and paleontology. Laboratory focuses on the identification and ecology of the avifauna of the Northeast, with field trips in eastern Massachusetts.

EEMB 5565. Lab for EEMB 5564. 1 Hour.

Accompanies EEMB 5564. Covers topics from the course through various experiments.

EEMB 5568. Wildlife Biology. 4 Hours.

Presents concepts and techniques utilized in the conservation and study of wild animals including the sociological aspects of management. Topics include habitat management, nonnative species, zoonoses, endangered species, legislation, and financing. Includes extended field trips to observe various ecosystems and wildlife.

EEMB 5569. Lab for EEMB 5568. 1 Hour.

Accompanies EEMB 5568. Covers topics from the course through various experiments.

EEMB 5589. Diving Research Methods. 2 Hours.

Presents experimental design, sampling methodology, statistical analysis, techniques, and the use of underwater equipment to conduct subtidal research.

EEMB 6402. Concepts and Trends in Evolution and Ecology. 4 Hours.

Presents key concepts and important recent advances in evolution and ecology, including interdisciplinary approaches to understanding the distributions, abundances, and diversity of species, organisms, and molecules. Topics include natural selection, adaptation, speciation, molecular evolution, global change, and perspectives on communities and ecosystems. Discusses and critiques current literature and methods.

EEMB 7100. Colloquium. 1 Hour.

Offers a seminar-style course that includes weekly lectures and presentations of selected topics. May be repeated up to four times.

EEMB 8507. Marine Biology Graduate Co-op Tutorial. 1 Hour.

Designed to complement learning during co-op. Offers students an opportunity to participate in activities to integrate academic learning and experiential learning including written reflections. Helps students share their experiences in the workplace through class discussions moderated by the instructor. May be repeated without limit.

EEMB 8674. Marine Biology Research Project. 1 Hour.

Offers an opportunity to design and implement a scientifically rigorous independent research project that builds upon current knowledge from the primary literature, under the supervision of a faculty advisor from the program. Students conduct research at any of the program’s locations and are then required to analyze data using rigorous statistical methods, write a journal-style research paper, and present their results in a research seminar.

EEMB 8960. Exam Preparation—Doctoral. 0 Hours.

Offers students an opportunity to prepare for the PhD qualifying exam under faculty supervision. May be repeated once.

EEMB 8982. Readings. 1-4 Hours.

Assigns students independent readings on selected topics in ecology, evolution, and marine biology. May be repeated without limit.

EEMB 8984. Research. 1-4 Hours.

Offers students an opportunity to conduct research. May be repeated without limit.

EEMB 8986. Research. 0 Hours.

Offers students an opportunity to conduct full-time research under faculty supervision. May be repeated without limit.

EEMB 9000. PhD Candidacy Achieved. 0 Hours.

Indicates successful completion of the doctoral comprehensive exam.

EEMB 9990. Dissertation. 0 Hours.

Offers theoretical and experimental research for the PhD degree. May be repeated once.

EEMB 9996. Dissertation Continuation. 0 Hours.

Offers dissertation supervision by members of the department. May be repeated without limit.