EEMB 1101. Foundations in Ecology and Evolutionary Biology. (4 Hours)

Introduces students to the foundational principles of ecology and evolutionary biology. Merges traditional lectures on foundational topics in ecology and evolutionary biology (adaptation, mechanisms of evolution, community and ecosystems ecology) with explorations of local field sites and an introduction to field ecology. Students spend several weeks of the semester designing and implementing independent field research projects, through which they are exposed to the foundation of scientific inquiry, including hypothesis testing, collecting, managing, and analyzing data, and presenting their findings.

Corequisite(s): EEMB 1102

Attribute(s): NUpath Natural/Designed World


EEMB 1102. Lab for EEMB 1101. (1 Hour)

Accompanies EEMB 1101. Covers topics from the course through various experiments.

Corequisite(s): EEMB 1101


EEMB 1105. Foundations in Ecological and Evolutionary Genomics. (4 Hours)

Introduces students to the foundational principles of molecular ecology with an emphasis on applications of high-throughput sequencing techniques to answer questions in ecology and evolutionary biology. Covers foundational topics in ecological and evolutionary genomics (central dogma, structure of nucleic acids, genetic variation, tools in molecular ecology, understanding genomes, and genomics). Practical skills development includes clean technique and proper bench skills; basic command line programming; understanding, quantifying, and analyzing sequence variation; and visualizing genomic data for formal scientific presentations.

Corequisite(s): EEMB 1106

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 1106. Lab for EEMB 1105. (1 Hour)

Accompanies EEMB 1105. Covers topics from the course through various experiments. Focuses on providing firsthand experience using tools from molecular ecology to test ecological and/or evolutionary hypotheses.

Corequisite(s): EEMB 1105


EEMB 1145. Beginning Scuba. (1 Hour)

Focuses on basic skin diving and scuba diving skills, with emphasis on safety. Requires lab fee. Requires ability to pass a swim test and basic comfort in the water.


EEMB 1450. Introduction to Marine Biology. (4 Hours)

Surveys the tremendous diversity of marine organisms in the context of the major marine ecosystems in which they are found. Explores interactions among organisms and how the physical and chemical environment influence marine organisms. Links changes on land to declines in organism numbers and diversity and explores the benefits humans gain from our relationship with the marine environment. Offers opportunities to investigate recent advances and understanding of marine organisms and their environments.

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 1990. Elective. (1-4 Hours)

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.


EEMB 2290. Ecology and Evolution of Behavior. (4 Hours)

Studies fundamental biological principles at behavioral, ecological, and evolutionary levels. Covers ethology, ecology, genetics, and comparative psychology, all within the conceptual framework of evolutionary theory. Explores both scientific practice and progress through readings, discussion, and projects. Illustrates the process by which biologists study questions about the evolutionary origin of behavior through a series of in-class activities, computer modeling assignments, interpretation of graphical data, collection and statistical analyses of behavioral data, as well as the generation and presentation of research. Does not focus on the neurological basis of behavior. Offers students an opportunity to become critical thinkers, critical readers, and to attain tools to interpret the world in a unique way. Requires permission of advisor.

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 2302. Ecology. (4 Hours)

Offers students an opportunity to learn about the environmental and biological processes that control the distribution and abundance of species and controlling factors that operate on individuals, populations, and communities. The lecture and laboratory introduce a set of generalizable concepts that are of fundamental importance to plant and animal life on the land and in the sea and provide hands-on experiential learning that reinforce concepts covered in lecture. Offers students an opportunity to become proficient in the following: (a) understanding research results the primary literature; (b) conducting a research experiment; (c) interpreting the results of in-class research; (d) communicating results as manuscript.

Prerequisite(s): ENGW 1111 with a minimum grade of C or ENGW 1102 with a minimum grade of C

Corequisite(s): EEMB 2303

Attribute(s): NUpath Formal/Quant Reasoning, NUpath Writing Intensive


EEMB 2303. Lab for EEMB 2302. (1 Hour)

Accompanies EEMB 2302. Covers topics from the course through various experiments.

Corequisite(s): EEMB 2302


EEMB 2400. Introduction to Evolution. (4 Hours)

Introduces evolutionary thinking, including contemporary examples of evolution. To understand the evolution of Charles Darwin’s “endless forms most beautiful,” the course adopts an integrative approach that includes information from ecology, genetics, molecular biology, biogeography, and paleobiology. Considers mechanisms of evolutionary change—how does it happen? Examines adaptation, the process by which attributes of an organism change to enhance fitness and the evolutionary history of life on our planet—what was the first living thing, how does speciation occur, what have we learned about evolution of life in the distant past, and how did humans evolve. Includes student presentations and analysis of scientific literature.

Prerequisite(s): BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or EEMB 1101 with a minimum grade of D- or ENVR 1400 with a minimum grade of D-

Attribute(s): NUpath Natural/Designed World


EEMB 2420. Fisheries Biology, Policy, and Conservation. (4 Hours)

Focuses on the study and management of economically valuable fish species. Studies the basic biology and ecology of fisheries species, quantifying and modeling their population biology to their interactions with each other and the environment. Requires students to read and analyze the scientific literature, to complete worksheets and writing assignments, and to develop and present research projects. Covers traditional stock assessment methods as well as how fisheries science and management has evolved more recently to integrate community- and ecosystem-level information. Reviews fisheries and how fishers are managed, their involvement in the management process, and the future fisheries in the United States and elsewhere.

Prerequisite(s): ENGW 1111 with a minimum grade of C or ENGW 1102 with a minimum grade of C

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World, NUpath Writing Intensive


EEMB 2700. Marine Biology. (4 Hours)

Examines biological aspects of natural ocean ecosystems and the physical processes that regulate them. Covers distributions, abundances, and interactions of marine organisms; interactions between organisms and the transformation and flux of energy and matter in marine ecosystems; and aspects of physiology related to marine species distributions, abundances, and roles. Students generate, evaluate, discuss, and present data from primary research and apply their knowledge of the scientific method and biological concepts through the creation of a written grant proposal.

Prerequisite(s): (BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or BIOL 1115 with a minimum grade of D- or EEMB 1101 with a minimum grade of D- ); (ENGL 1111 with a minimum grade of C or ENGL 1102 with a minimum grade of C or ENGW 1111 with a minimum grade of C or ENGW 1102 with a minimum grade of C )

Corequisite(s): EEMB 2701

Attribute(s): NUpath Natural/Designed World, NUpath Writing Intensive


EEMB 2701. Lab for EEMB 2700. (1 Hour)

Accompanies EEMB 2700. Covers topics from the lecture course through discussions and experiments.

Corequisite(s): EEMB 2700


EEMB 2990. Elective. (1-4 Hours)

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.


EEMB 3001. Genetics and Evolution in Action. (4 Hours)

Uses current topics in genetics and evolution to study how evolution shapes the world around us and impacts our society. Explores cutting-edge genetic approaches and concepts using case studies, media reports, and scientific papers. Covers emerging topics such as consumer genetic testing of human ancestry and genetic risk, tests and analyses of novel viruses such as Covid19, the evolution of antibiotic resistance, origins of immunity genetics of domestication, and human microbial interactions. Includes discussions and writing exercises designed to offer students an opportunity to develop critical scientific thinking and problem-solving skills.

Prerequisite(s): BIOL 2301 with a minimum grade of D-


EEMB 3450. Physiological Adaptations to the Environment. (4 Hours)

Explores the evolutionary mechanisms by which organisms adapt physiologically to survive, and thrive, in diverse, often seemingly “hostile,” habitats. Examines paleo- and modern examples of adaptation with the goal of predicting species success or failure as our planetary environment changes rapidly. Topics include adaptation of cellular metabolism, adaptations to variable oxygen availability and to changes in pH, the roles of water and microsolutes in regulation of the internal environment of cells, and the effects of temperature on cellular function and the biogeographic distribution of organisms. Includes student presentations and analysis of scientific literature. Requires junior or senior standing; sophomores admitted by permission of instructor; EEMB 2400 or ENVR 2400 recommended but not required.

Prerequisite(s): (EEMB 1105 with a minimum grade of D- or BIOL 1113 with a minimum grade of D- or BIOL 2299 with a minimum grade of D- ); (CHEM 1161 with a minimum grade of D- or CHEM 1214 with a minimum grade of D- )

Attribute(s): NUpath Natural/Designed World


EEMB 3455. Ecosystems Ecology. (4 Hours)

Focuses on the foundational principles of ecosystems ecology. Examines the flow of energy and materials through both the biosphere (plants, animals, and microbes) and the geosphere (soils, atmospheres, and oceans) and the role that humans are playing in altering these key fluxes. Studies elemental cycles that are critically important for human and environmental sustainability—including carbon, nitrogen, and phosphorus—and examines similarities and differences in these cycles and flows while drawing on examples from both terrestrial and marine systems. Seeks to understand how changes in ecosystem structure ultimately affect ecosystem function and how this translates into the important services ecosystems provide.

Prerequisite(s): BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or CHEM 1161 with a minimum grade of D- or CHEM 1214 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or ENVR 1200 with a minimum grade of D- or ENVR 2200 with a minimum grade of D-

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 3460. Conservation Biology. (4 Hours)

Explores conservation biology, an interdisciplinary science that focuses on conservation of biological diversity at multiple levels. Emphasizes the causes and consequences of biodiversity loss and demonstrates how ecological and evolutionary principles are applied to conservation problems. Covers sustainability; climate change; introduced species; conservation of threatened and endangered species; and pollution, disease, and habitat restoration using examples from marine, aquatic, and terrestrial systems. Offers students an opportunity to read, discuss, evaluate, and present data from primary research through written assignments and oral debates and to apply this knowledge to conservation issues. Emphasizes critical thinking, problem solving, and recognizing multiple perspectives.

Prerequisite(s): BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or BIOL 1115 with a minimum grade of D- or EEMB 1101 with a minimum grade of D- or ENVR 1101 with a minimum grade of D- or ENVR 1400 with a minimum grade of D-

Attribute(s): NUpath Writing Intensive


EEMB 3465. Ecological and Conservation Genomics. (4 Hours)

Offers an overview of ecological and conservation genetics, an interdisciplinary science that focuses on understanding the processes that determine genetic diversity at the individual to population level. Focuses on fundamental concepts in evolutionary ecology and population and quantitative genetics, then applies those concepts to solving real-world problems in conservation science. Covers harvested populations, inbreeding, climate change, introduced species, conservation of threatened and endangered species, adaptation, and habitat restoration. Exposes students to multiple sides of these issues and the science that underpins them. Offers students an opportunity to develop the R programming skills required to analyze the complex data sets that often emerge when addressing cutting-edge questions in genetics. Includes writing and coding exercises and mathematical derivations. Emphasizes critical thinking and problem solving.

Prerequisite(s): BIOL 2301 with a minimum grade of C- or CS 2500 with a minimum grade of C- or ECON 2350 with a minimum grade of C- or EEMB 2400 with a minimum grade of C- or ENVR 2500 with a minimum grade of C- or MATH 3081 with a minimum grade of C-

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 3466. Disease Ecology. (4 Hours)

Covers the fundamentals of disease ecology and evolution. Focuses on how disease can impact the physiology of organisms and how this can, in turn, alter communities and ecosystems. Topics include mathematical theory on host-pathogen interactions; empirical studies of human, wildlife, insect, and plant host populations; emerging infectious diseases; effects on host behavior; host-parasite coevolution; multihost and multipathogen systems; and anthropogenic effects on disease. Includes writing exercises, with a special emphasis on critical thinking and problem solving.

Prerequisite(s): BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or EEMB 1101 with a minimum grade of D- or ENVR 1101 with a minimum grade of D- or ENVR 1400 with a minimum grade of D-

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 3470. Coastal Ecology and Sustainability. (4 Hours)

Designed to provide an integrated exposure to issues surrounding the ecology and sustainability of coastal and estuarine systems, with a particular focus on urban harbors. Exposes students to both the diversity and complexity of coastal habitats that exist both locally (salt marshes and seagrass beds) and globally (mangroves) and the mechanisms of estuarine and coastal functioning (geomorphology, biogeochemistry, microbial ecology, food webs, fisheries). Considers the ecosystem services provided by coastal systems and how those services are altered through human pressures.

Prerequisite(s): (CHEM 1211 with a minimum grade of D- or CHEM 1151 with a minimum grade of D- ); (EEMB 1101 with a minimum grade of D- or BIOL 1107 with a minimum grade of D- or BIOL 1111 with a minimum grade of D- or BIOL 1115 with a minimum grade of D- )


EEMB 3475. Wildlife Ecology. (4 Hours)

Focuses on wildlife ecology and management, with an emphasis on terrestrial species. Introduces habitat use, behavior, wildlife conservation, parasites and pathogens, wildlife sampling, wildlife management, food and nutrition, population viability, and conservation genetics. Offers students an opportunity to engage in analyzing primary literature, collection, interpretation, and wildlife data and using basic mathematical models.

Prerequisite(s): EEMB 2302 with a minimum grade of C

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 3555. Networks and Natural Systems. (4 Hours)

Covers the properties of diverse biological networks and explores foundational computational methods for analyzing, visualizing, and performing statistical investigations of networked data. From social networks and cities to ecosystems and evolution, methods from network science provide powerful tools for understanding and investigating the natural and modern world. Moving beyond description, a key objective of the course is to synthesize the diversity of biological networks and investigate how scientists have uncovered remarkable regularities in networked systems by applying approaches from scaling theory to biological networks. Based on a series of case-studies, focuses on how to elucidate the structure and function of biological networks using empirical data. Requires scientific programming.

Prerequisite(s): ENVR 2500 with a minimum grade of D or DS 2000 with a minimum grade of D or CS 2500 with a minimum grade of D or MATH 2280 with a minimum grade of D or PHYS 1211 with a minimum grade of D or PHTH 2210 with a minimum grade of D or PSYC 2320 with a minimum grade of D

Attribute(s): NUpath Analyzing/Using Data, NUpath Natural/Designed World


EEMB 3700. Desert Ecology. (4 Hours)

Offers students an opportunity to obtain a basic understanding of fundamental ecological processes taking place in desert environments. Familiarizes students with how environmental and biological processes interact and influence the distribution and abundance of species in these arid biomes while recognizing the impact that human societies have on desert life and identifying sustainable solutions to ameliorate our ecological footprint. Introduces students to foundational concepts of fundamental importance to desert plant and animal life. Uses an ecological perspective to surround students with a rich social/cultural milieu including interactions with Israeli, Palestinian, and Bedouin communities.

Prerequisite(s): EEMB 2302 with a minimum grade of D- ; EEMB 2303 with a minimum grade of D-

Attribute(s): NUpath Difference/Diversity


EEMB 3990. Elective. (1-4 Hours)

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.


EEMB 4000. Applied Conservation Biology. (4 Hours)

Studies landscape-scale conservation in Transylvania and the Carpathian Mountains of Romania. Working intensively with Foundation Conservation Carpathia, explores efforts to build Europe’s largest national park. Offers students an opportunity to learn from local conservation leaders, collect data, and develop plans to help launch the “Yellowstone of Europe.” Focuses on large carnivore conservation (brown bears, lynx, and wolves); sustainable agriculture; resource management in a country formerly under communist rule; and balancing urban and rural conservation needs. Explores Romania’s rich cultural heritage in Sighisoara, a UNESCO World Heritage Site, and Vacaresti Nature Park, a constructed urban wetland in the heart of Bucharest. Requires prior completion of one laboratory science course or permission of instructor.

Attribute(s): NUpath Integration Experience, NUpath Natural/Designed World


EEMB 4001. Landscape and Restoration Ecology. (4 Hours)

Topics include ecosystem processes, spatial patterns, disturbance, species distributions, invasive species, and habitat loss. Offers students an opportunity to participate in activities in which they look at and interpret spatial data. Course format includes group work, analyzing the scientific literature, and in-class activities.

Prerequisite(s): BIOL 1107 with a minimum grade of C or BIOL 1111 with a minimum grade of C or EEMB 1101 with a minimum grade of C or ENVR 1101 with a minimum grade of C or ENVR 1400 with a minimum grade of C

Attribute(s): NUpath Natural/Designed World


EEMB 4010. Biology of Mammals. (4 Hours)

Surveys the mammals of the world, including their evolution, morphology, physiology, behavior, and ecology. Students conduct a research project in which they investigate the morphology, evolution, ecology, and behavior of a species and present their findings to the class. Includes reading and analyzing the scientific literature and conducting in-class activities.

Prerequisite(s): BIOL 1113 with a minimum grade of D- or BIOL 2299 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or EEMB 2400 with a minimum grade of D-

Attribute(s): NUpath Natural/Designed World


EEMB 4548. Sociobiology. (4 Hours)

Studies sociobiology, a field of biology that strives to understand the biological basis of social behavior in animals. Sociobiology is a multidisciplinary science, meshing together ethology (animal behavior), ecology, genetics, population biology, and comparative psychology, all within the conceptual framework of evolutionary theory. Why do animals live in societies? Why do animals cooperate and sometimes show extreme forms of altruism? What are the costs and benefits of group living? Reviews studies on nonhuman animals that demonstrate sociobiological principles by using a series of in-class activities, computer-modeling assignments, interpretation of graphical and tabulated data, collection and statistical analyses of behavioral data, as well as the generation and presentation of research.

Prerequisite(s): BIOL 2301 with a minimum grade of D-

Attribute(s): NUpath Analyzing/Using Data


EEMB 4990. Elective. (1-4 Hours)

Offers elective credit for courses taken at other academic institutions. May be repeated without limit.


EEMB 4992. Directed Study. (1-4 Hours)

Offers independent work under the direction of members of the department on a chosen topic. Course content depends on instructor. May be repeated without limit.


EEMB 5130. Ecological Dynamics. (4 Hours)

Offers a comprehensive overview of mathematical and computational concepts needed to construct (meta)population, (meta)community, and (meta)ecosystem models. Focuses on how to mathematically derive and model processes (growth, trophic and nontrophic species interactions, dispersal, and environmental variability) to understand patterns of population abundance and species diversity. Emphasizes the mathematical tools required to analyze the dynamical behavior of ecological models (stability, invasion, graphical, and numerical analyses) and validate model predictions using empirical data (via maximum likelihood and optimization methods). Sophomores admitted by permission of instructor.

Prerequisite(s): ((MATH 1241 (may be taken concurrently) with a minimum grade of D- or MATH 1251 (may be taken concurrently) with a minimum grade of D- or MATH 1341 (may be taken concurrently) with a minimum grade of D- ); (ENGW 1111 with a minimum grade of C or ENGW 1102 with a minimum grade of C or ENGL 1111 with a minimum grade of C or ENGL 1102 with a minimum grade of C )) or graduate program admission

Corequisite(s): EEMB 5131

Attribute(s): NUpath Writing Intensive


EEMB 5131. Lab for EEMB 5130. (1 Hour)

Accompanies EEMB 5130. Offers supervised lab sessions designed to show how the topics covered in the lectures can be addressed in industry-standard programming environments.

Corequisite(s): EEMB 5130


EEMB 5303. Marine Biology Careers Seminar. (1 Hour)

Covers the information and tools needed to begin pursuing career opportunities in marine biology. Encourages students to explore a variety of career paths, construct résumés, contact potential employers for their internship and permanent positions. Presents invited speakers from state and federal agencies, and from private consulting firms, to talk about their work and career track.


EEMB 5305. Professional Development for Ocean Sciences. (2 Hours)

Designed to assist Three Seas students in securing a graduate research internship. Seeks to provide students with the information needed to pursue diverse career opportunities in marine biology. Provides hands-on experience with an array of science communication tools, including resumés/CVs, cover letters, and social media. Speakers from academia, informal science education organizations, and the media present talks on their work and career tracks.


EEMB 5504. Biology of Corals. (2 Hours)

Covers a variety of topics including basic coral biology, the coral/algal symbioses, the mechanisms of coral bleaching, coral microbiology and disease, coral calcification and ocean acidification, and coral speciation and hybridization. Supplements lectures with readings from the primary literature. Focuses on active areas of research and hands-on learning through lab and field activities.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Corequisite(s): EEMB 5505

Attribute(s): NUpath Analyzing/Using Data


EEMB 5505. Lab for EEMB 5504. (1 Hour)

Accompanies EEMB 5504. Focuses on relevant research questions while providing practical training techniques in coral biology. Hands-on learning includes visual surveys of reef transects, quadrat sampling, coral identification, enumeration of zooxanthellae with a hemocytometer, PAM fluorometry, ImageJ analysis, coral homogenization, and Vibrio plating.

Corequisite(s): EEMB 5504


EEMB 5506. Biology and Ecology of Fishes. (2 Hours)

Covers fundamental concepts in reef fish biology, ecology, and conservation. Additional lecture coursework includes analysis of both group and individual research projects conducted in lab. Presents recent or ongoing research projects by the instructor and guest lecturers. Discussions are based on papers from the scientific literature and relate topics about processes and patterns of fish recruitment, reproduction, dispersal, evolution, conservation, and management.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Corequisite(s): EEMB 5507


EEMB 5507. Lab for EEMB 5506. (1 Hour)

Accompanies EEMB 5506. Studies methods to conduct research on reef fishes through class exercises and individual research projects. Hands-on learning includes common fish transect methodology, reef fish identification, cast net and handline (on scuba) fishing techniques, and specimen dissection. Emphasizes analyzing and presenting the data and writing clearly and effectively about scientific research through the lab reports.

Corequisite(s): EEMB 5506


EEMB 5508. Marine Birds and Mammals. (3 Hours)

Studies principles of classification, anatomy, physiology, behavior, and evolution of seabirds and marine mammals. Also addresses conservation and protection of animals and essential habitat. Includes field trips to observe local species.

Prerequisite(s): (BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- ) or graduate program admission


EEMB 5510. New England Marine Biomes. (4 Hours)

Investigates the major biomes in the northwest Atlantic, including their habitats—rocky intertidal, tidal estuaries, seagrass beds, kelp forest/rocky reef complex, soft sediments, salt marshes, and continental shelf. Studies the major chemical, physical, geological, and biological forces that shape each habitat. Investigates the ecological framework of each habitat, both in the field and in hands-on exercises. Examines the adaptations of plants, algae, and animals to their respective ecosystem. Offers students an opportunity to develop an appreciation for human-induced changes in each habitat and biome and the conservation and restoration efforts currently being used.

Prerequisite(s): (EEMB 2302 with a minimum grade of D- or EEMB 2700 with a minimum grade of D- or EEMB 3460 with a minimum grade of D- or EEMB 3475 with a minimum grade of D- or EEMB 4001 with a minimum grade of D- ) or graduate program admission


EEMB 5512. Tropical Terrestrial Ecology. (1 Hour)

Studies the animals, plants, and ecosystems of the new world tropics, with the community structure and diversity of terrestrial Jamaican habitats as an example. Includes field trips to lowland forests, carbonate caves, and the Blue Mountain mist-montane forest. The issue of land use and development vs. conservation is a recurring theme.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission


EEMB 5516. Oceanography. (4 Hours)

Offers an integrated overview of physical, chemical, biological, and geological processes operating in the world ocean. Seemingly unrelated topics like plate tectonics, oscillating currents and waves in the atmosphere, the activities of microbes and phytoplankton, and land-use practices in the middle of the continent have global reach and interact with each other in surprising yet understandable ways. Examines how new technologies have allowed stunning insights into global weather and climate, the deep sea, biodiversity, and how the biogeochemistry of the oceans can be measured and understood. Presents data use and analysis and formal reasoning used in marine science. Views the ocean as a “system of systems” where integration of experience from disparate disciplines is key.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Corequisite(s): EEMB 5517

Attribute(s): NUpath Analyzing/Using Data, NUpath Formal/Quant Reasoning


EEMB 5517. Lab for EEMB 5516. (1 Hour)

Accompanies EEMB 5516. Offers experiential field and laboratory exercises in oceanography. The New England rocky intertidal, subtidal, wetlands, barrier islands, and dunes provide opportunities for field exercises in marine geology, physical oceanography, and marine ecology. Investigates processes affecting changes in the global ocean, such as ocean acidification; temperature stress in organisms; hydrodynamic drag and lift; suspension feeding; and the ecophysiology of reef corals, boreal invertebrates, and macroalgae.

Corequisite(s): EEMB 5516


EEMB 5518. Ocean and Coastal Processes. (2 Hours)

Examines the coupling between physical and biological processes on coral reefs and adjacent habitats. Focuses on biophysical, oceanographic, and benthic-pelagic processes acting in coral reef and associated nearshore ecosystems. Specific topics include oceanographic forcing mechanisms, organismal biomechanics, hydrodynamics, and nutrient dynamics.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Corequisite(s): EEMB 5519


EEMB 5519. Lab for EEMB 5518. (1 Hour)

Accompanies EEMB 5518. Studies techniques such as estimation of turbulent diffusion coefficients, mean speed, and logarithmic layer parameters using analysis of video imagery (NIH ImageJ); deploying and recovering zooplankton traps and nets and estimating mortality using vital dyes; use of GoPro for behavioral analyses (sediment shedding in corals); estimating flux rates from active suspension feeders like sponges using dye release; and use of the YSI Exo Sonde to measure different water quality parameters.

Corequisite(s): EEMB 5518


EEMB 5520. Tropical Marine Ecology. (2 Hours)

Highlights and explores the ecological characteristics and current threats facing four tropical ecosystems—coral reefs, seagrass beds, mangrove forests, and tropical lowland rain forests. Explores the connectivity between these ecosystems and the services each provides. Examines how these ecosystems have changed under past threats and are projected to change in future conditions. Includes formal lectures, informal lectures provided in the field, field demonstrations, and interpretive hikes.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission


EEMB 5522. Experimental Design Marine Ecology. (4 Hours)

Includes introduction to and application of observational methods in three local marine habitats, experimental design, statistical analysis, R statistical computing and graphics software, and principles of marine ecology. Combines lecture, hand-on research experience, and computer laboratory and includes reading and analyzing the scientific literature and developing research projects. At the end of the semester, students are expected to demonstrate an integrative mastery of course topics by writing a scientific manuscript about a class experiment. Seeks to prepare students for practicing ecology in new environments and to provide students with the foundational knowledge necessary for pursuing more complex concepts in experimental design, statistical analysis, and marine ecology.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Attribute(s): NUpath Analyzing/Using Data, NUpath Writing Intensive


EEMB 5525. Advanced Field Methods in Marine Ecology. (3 Hours)

Explores the methods used to build and complete scientific studies in marine ecology from observation to data analysis and interpretation within the context of the northwest Atlantic Ocean. Offers students an opportunity to build quantitative skills by understanding how and when to apply different statistical methods to a range of ecological datasets. Studies how to appropriately interpret results and effectively communicate the interpretation to any audience. Applies these skills to additional study systems outside the marine environment of the northwest Atlantic.

Prerequisite(s): ENVR 2500 with a minimum grade of D- or graduate program admission


EEMB 5528. Marine Conservation Biology. (3 Hours)

Examines several critical issues facing marine ecosystems, including invasive species, marine pollution and eutrophication, fisheries impacts, physical alteration of habitats, and global climate change. Offers students an opportunity to spend field time surveying intertidal and subtidal habitats within the San Juan Islands and Friday Harbor Marine Reserve and to conduct independent research projects.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission

Attribute(s): NUpath Analyzing/Using Data, NUpath Ethical Reasoning


EEMB 5532. Physiological and Molecular Marine Ecology. (3 Hours)

Explores the physiological responses of marine organisms to variations in environmental factors. Uses complementary techniques, including molecular and physiological approaches, to determine genetic relationships at the species and population level and elucidate the mechanistic basis of organismic responses to environmental conditions at the level of genes and gene products.

Prerequisite(s): BIOL 2311 with a minimum grade of D- or EEMB 2302 with a minimum grade of D- or graduate program admission


EEMB 5533. Marine Invertebrate Zoology and Botany. (2 Hours)

Explores major groups of marine macroalgae and marine invertebrates, their ecological roles, and interrelationships. Identifies defining features of these groups and the evolutionary and ecological drivers leading to adaptations. Emphasizes important groups in the Pacific Northwest habitats, including kelp forest and rocky intertidal habitats. Hands-on learning with corequisite lab includes field identification; visits to intertidal and subtidal marine environments; and specimen observation, dissection, preparation, and cataloging.

Corequisite(s): EEMB 5535


EEMB 5535. Lab for EEMB 5533. (1 Hour)

Accompanies EEMB 5533. Covers topics from the course through various experiments.

Corequisite(s): EEMB 5533


EEMB 5538. Conservation and Restoration of Marine Systems. (3 Hours)

Designed to foster an understanding of conservation and restoration strategies in the nearshore marine system using a real-world case study approach. Students collect and analyze historical and newly collected data and use this to inform a conservation and restoration plan within the context of local, state, and federal laws. Uses real-world examples (such as aquaculture, seagrass restoration, and shoreline hardening) to build ecological goals. Seeks to build understanding and appreciation of the input of all stakeholders, including that of marginalized groups. Offers students an opportunity to produce a high-level and annotated deliverable that could serve as a template for real-world use.


EEMB 5540. Changing Global Oceans. (2 Hours)

Investigates the major drivers to short-, medium-, and long-term changes in the world’s oceans. Compares the role of natural and human-induced changes in ocean systems. Key areas focus on the role of nonhuman animals in modifying and mitigating oceanic and atmospheric change. Explores the linkages among oceans and atmosphere through examples in the Pacific Northwest and worldwide.

Corequisite(s): EEMB 5541


EEMB 5541. Lab for EEMB 5540. (1 Hour)

Accompanies EEMB 5540. Students participate in daily, topical, paper discussions, carry out laboratory exercises, or explore and collect data in field exercises. Offers students an opportunity to acquire skills such as using oceanographic equipment, modeling simulations, data collection, and data analysis.

Corequisite(s): EEMB 5540


EEMB 5542. Marine Spatial Planning. (4 Hours)

Investigates issues of marine and coastal spatial planning (MCSP) that include offshore wind power siting, fisheries and aquaculture management, natural resource extraction, marine mammal conservation, and/or living shoreline protection and mitigation. Covers the spatial planning process from question to deliverable strategy, including assessment of stakeholder needs and potential ecosystem impacts. Offers students an opportunity to acquire and assess data, apply appropriate statistical tools, and develop spatial maps using geographic information systems (GIS) and other software. Also covers how to synthesize the planning process and develop and evaluate recommendations.

Prerequisite(s): EEMB 2302 with a minimum grade of D- or EEMB 2700 with a minimum grade of D- or EEMB 3460 with a minimum grade of D- or EEMB 4001 with a minimum grade of D-

Attribute(s): NUpath Capstone Experience


EEMB 5546. Sustainability of the Land-Sea Interface. (3 Hours)

Explores the current issues facing management and conservation of the land-sea interface, also known as the coastal transition zone (CTZ). Evaluates the mitigation, conservation, and restoration tools that are applied to human use of the land-sea interface. Observes these tools during site visits and discusses strategies with experts in sustainability of these habitats. Synthesizes the scientific literature on CTZ tools in the northwest Atlantic and other regions with pressing sustainable land-sea use issues. Offers students an opportunity to develop skills in prioritizing and advocating for particular conservation strategies and to practice science communication skills to effectively reach a broad audience.

Prerequisite(s): (EEMB 2302 with a minimum grade of D- or EEMB 2700 with a minimum grade of D- or EEMB 3460 with a minimum grade of D- or EEMB 4001 with a minimum grade of D- ) or graduate program admission

Attribute(s): NUpath Writing Intensive


EEMB 5589. Diving Research Methods. (2 Hours)

Presents experimental design, sampling methodology, statistical analysis, techniques, and the use of underwater equipment to conduct subtidal research.


EEMB 6465. Ecological and Conservation Genomics. (4 Hours)

Provides an overview of ecological and evolutionary genomics. Covers foundational mathematical concepts in population in quantitative genetics, from individual loci up to whole genomes. Concepts covered include Hardy-Weinberg equilibrium, F statistics, signatures of natural selection in genomes and methods for detecting them, analysis of quantitative genetic evolution, hybridization, and gene expression. Also covers modern statistical methods used to analyze genomic data using the free and open source R programming environment. Builds knowledge through reading of the primary literature and advanced problem sets. The final project requires students to complete a novel data analysis of an open source genomics data set and write a research paper.


EEMB 6470. Coastal Ecology and Sustainability. (4 Hours)

Offers an integrated exposure to issues surrounding the ecology and sustainability of coastal marine systems, with a particular focus on urban harbors. Exposes students to both the diversity and complexity of coastal habitats that exist locally (salt marshes and sea grass beds) and globally (mangroves) and the mechanisms of estuarine and coastal ecosystem functioning (geomorphology, biogeochemistry, microbial ecology, food webs, fisheries). Throughout the course, students focus on translating the basic science of coastal ecology into relevant sustainability practices.


EEMB 6475. Advanced Wildlife Ecology. (4 Hours)

Focuses on wildlife ecology and management, with an emphasis on terrestrial species. Covers habitat use, behavior, wildlife conservation, parasites and pathogens, wildlife sampling, wildlife management, food and nutrition, population viability, and conservation genetics. Engages students in analyzing primary literature and wildlife data, collection, interpretation, and using basic mathematical models.


EEMB 7101. Seminar in Marine Sciences. (2 Hours)

Offers students an opportunity to lead critical discussions of recent and classic papers from primary and secondary literature in marine sciences. Discusses the important scientific paradigms and addresses strengths and weaknesses of these papers from the perspective of scientific communication, including design and presentation of data in figures and tables; the role of synthesis in justifying new concepts; and how terminology and jargon evolve in scientific subdisciplines. Students write occasional reviews of these papers as if they had just been submitted to a journal for consideration.


EEMB 7102. Seminar in Ecology and Evolutionary Biology. (2 Hours)

Offers an overview of major concepts in the fields of ecology and evolution and how these concepts can be synthesized under a common framework. The first half of the course is organized according to major areas of evolutionary biology, from quantitative genetics to population genetics and phylogenetics and their synthesis. Quantitative genetics, population genetics, and phylogenetics have been historically separate fields and have only recently been synthesized through genomics. Note that quantitative genetics is a field that studies the evolution of phenotypes and requires no genetic information. The second half of the course introduces major concepts in ecology and is designed to introduce students to the major historical underpinnings of community ecology so as to understand the utility (or lack thereof) of these concepts for modern ecology.


EEMB 7103. Seminar in Sustainability Sciences. (2 Hours)

Explores key papers that have shaped modern theory, methodologies, and practices of sustainability science. Sustainability science hinges on integrating social and ecological sciences to assess the sustainability of human-environment interactions. From the social science dimension, many past studies focused on understanding how values, beliefs, and social norms shape human behavior. From an ecological perspective, much work focused on the influence of various institutional arrangements on resource and environmental sustainability. Importantly, a coupled natural-human or social-ecological systems (SES) perspective focuses on the inherently dynamic nature of these systems and interactions.


EEMB 7104. Seminar in Geosciences. (2 Hours)

Exposes graduate students pursuing a PhD in marine and environmental sciences to classical and recent high-impact papers in the fields of recent and deep earth history, landform evolution, microbes and their role in global biogeochemical cycling, nutrient stoichiometry, the global carbon cycle, geochemical proxies, evolution of ocean chemistry, oceanic acidification, the role of organisms in sediment and rock production, and geochemical paleoproxies. Examines applications of the above disciplines to mitigating the impacts of anthropogenic impacts on the Earth system. This is a guided readings course.


EEMB 7674. Marine Biology Research Project. (1 Hour)

Offers an opportunity to design and implement a scientifically rigorous independent research project that builds upon current knowledge from the primary literature, under the supervision of a faculty advisor from the program. Students conduct research at any of the program’s locations and are then required to analyze data using rigorous statistical methods, write a journal-style research paper, and present their results in a research seminar.


EEMB 8101. Readings in Marine Sciences. (2 Hours)

Designed to prepare PhD students with a concentration in marine sciences for a career in their field by offering an opportunity to learn fundamental aspects of the discipline through readings. Each student works with their Northeastern committee members at their first committee meeting to identify one reading topic per committee member. Committee members provide guidance for the student’s readings around their topic. Students meet with each committee member throughout the semester to discuss the readings, ask questions, and clarify any aspects of their topics.


EEMB 8102. Readings in Ecology and Evolutionary Biology. (2 Hours)

Designed to prepare PhD students with a concentration in ecology and evolutionary biology for a career in their field by offering an opportunity to learn fundamental aspects of the discipline through readings. Each student works with their Northeastern committee members at their first committee meeting to identify one reading topic per committee member. Committee members provide guidance for the student’s readings around their topic. Students meet with each committee member throughout the semester to discuss the readings, ask questions, and clarify any aspects of their topics.


EEMB 8103. Readings in Sustainability Sciences. (2 Hours)

Designed to prepare PhD students with a concentration in sustainability for a career in their field by offering an opportunity to learn fundamental aspects of the discipline through readings. Each student works with their Northeastern committee members at their first committee meeting to identify one reading topic per committee member. Committee members provide guidance for the student’s readings around their topic. Students meet with each committee member throughout the semester to discuss the readings, ask questions, and clarify any aspects of their topics.


EEMB 8104. Readings in Geosciences. (2 Hours)

Designed to prepare PhD students with a concentration in geosciences for a career in their field by offering an opportunity to learn fundamental aspects of the discipline through readings. Each student works with their Northeastern committee members at their first committee meeting to identify one reading topic per committee member. Committee members provide guidance for the student’s readings around their topic. Students meet with each committee member throughout the semester to discuss the readings, ask questions, and clarify any aspects of their topics.


EEMB 8982. Readings. (1-4 Hours)

Assigns students independent readings on selected topics in ecology, evolution, and marine biology. May be repeated without limit.


EEMB 8984. Research. (1-4 Hours)

Offers students an opportunity to conduct research. May be repeated without limit.


EEMB 8986. Research. (0 Hours)

Offers students an opportunity to conduct full-time research under faculty supervision. May be repeated without limit.


EEMB 9000. PhD Candidacy Achieved. (0 Hours)

Indicates successful completion of the doctoral comprehensive exam.


EEMB 9990. Dissertation Term 1. (0 Hours)

Offers theoretical and experimental research for the PhD degree.

Prerequisite(s): EEMB 9000 with a minimum grade of S


EEMB 9991. Dissertation Term 2. (0 Hours)

Offers dissertation supervision by members of the department.

Prerequisite(s): EEMB 9990 with a minimum grade of S


EEMB 9996. Dissertation Continuation. (0 Hours)

Offers dissertation supervision by members of the department.

Prerequisite(s): EEMB 9991 with a minimum grade of S or Dissertation Check with a score of REQ