Marine and Environmental Sciences

Website

Geoffrey C. Trussell, PhD
Professor and Chair

Marine Science Center
781.581.7370
781.581.6076 (fax)
gradmes@northeastern.edu

Graduate Coordinator
Jonathan Grabowski, PhD, Associate Professor, j.grabowski@northeastern.edu

The Department of Marine and Environmental Sciences graduate program offerings include core capacities in marine biology, ecology, and evolution. Students benefit from top-notch research facilities at both the Marine Science Center and the main campus in Boston. The MS in Marine Biology program seeks to prepare students for entry- and mid-level careers in marine research. The doctoral program in ecology, evolution, and marine biology is designed to prepare graduates for careers in academia, government agencies, and the private sector.

Earth and Environmental Sciences Courses

ENVR 5105. Geophysics. 4 Hours.

Examines the physical processes of sediment erosion, transportation, and deposition and the origin of sediment. Emphasis is on the effect of coastal marine processes and resultant responses of the coast. Topics include the dynamics of waves and currents and such coastal landforms as beaches, barriers, salt marshes, and bluffed and rocky coasts. (a) ENVR 1112, ENVR 1200, or graduate standing and (b) MATH 1241, MATH 1251, MATH 1341, or graduate standing and (c) junior, senior, or graduate standing.

ENVR 5110. Coastal Sedimentation. 4 Hours.

Examines a current environmental issue or topic through an understanding of the scientific principles controlling the process, review of alternative actions, and inquiry into societal implications of the issue. Topics include groundwater supply, groundwater contamination, coastal erosion and flooding, or impacts of land development. Prereq. Junior, senior, or graduate standing.

ENVR 5115. Advanced Topics in Environmental Geology. 4 Hours.

Examines selected topics in geology through an understanding of the basic processes, materials, and evolution. Topics include basin analysis, landform evolution, volcanology, or regional geology. Prereq. Junior, senior, or graduate standing.

ENVR 5120. Advanced Topics in Geology. 4 Hours.

Introduces spatial data analysis through geographical information system (GIS) systems. Topics include basics of cartography, cartographic transformations on the computer, data input, data sorting and presentation, and statistical analysis. Emphasis is on practical applications of GIS methods. Prereq. Junior, senior, or graduate standing.

ENVR 5190. Soil Science. 4 Hours.

Provides a description and evaluation of the physical, chemical, and biological properties of soils. Includes soil formation, soil types, and processes that occur in soil including the importance of these processes for the soil productivity and management of soil. Also covers sources, reactions, transports, and fates of chemical species in soils and associated water and air environments, as well as the chemical behavior of elements and compounds and the phenomena affecting natural and anthropogenic materials in soils. Prereq. (a) ENVR 1112, ENVR 1200, ENVR 2310, or graduate standing and (b) junior, senior, or graduate standing.

ENVR 5200. Geology Seminar. 4 Hours.

Offers an analysis of selected topics in geology for advanced study. Topics are selected from current areas of active research in the field. Prereq. Junior, senior, or graduate standing.

ENVR 5201. Geologic Field Seminar. 4 Hours.

Studies aspects of geology/environmental science associated with a particular field setting, in the classroom, followed by an intensive field investigation. Examples include carbonate petrology and reef ecology, then field studies in the Bahamas; glacial geology and volcanology, followed by field studies in Iceland; or stratigraphy of the U.S. Southwest, with field studies in the Grand Canyon. Focuses on using field observations and field data to interpret modern and ancient geologic processes.

ENVR 5202. Environmental Science Field Seminar Abroad. 4 Hours.

Offers an intensive environmental science field study experience associated with a particular off-campus geographic setting, such as Iceland, Newfoundland, Bahamas, etc. Offers students an opportunity to learn the principles of field study, to learn to recognize and record significant data, and to reach conclusions about a range of field-based problems being studied.

ENVR 5210. Environmental Planning. 4 Hours.

Examines aspects of surface runoff from geomorphic and hydrologic perspectives. Develops methods for description and calculation of major river and drainage basin processes and applies the results to the planning process. Examines human modification of these systems—including urbanization, dams, and channelization—and applies this information to an understanding of regulatory processes. This is a writing-intensive course. Prereq. (a) Junior or senior standing and ENGW 1111, ENGW 1102, ENGL 1111, or ENGL 1102, or (b) graduate standing.

ENVR 5230. Structural Geology. 4 Hours.

Focuses on the description and origin of rock structures, with emphasis on interpretation of the mechanics of deformation. Lab analyses of structural features and problems utilize geologic maps, structural models, stereograms, petrographic microscope, rock specimens, and field exercises. Prereq. (a) ENVR 1200, ENVR 1201, and junior or senior standing or (b) graduate standing. Coreq. ENVR 5231.

ENVR 5231. Lab for ENVR 5230. 1 Hour.

Accompanies ENVR 5230. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5230.

ENVR 5240. Sedimentary Basin Analysis. 4 Hours.

Presents the analysis of sedimentary basins based on detailed study of sedimentary petrology, sedimentary structures, and stratigraphic sequences and fossils. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5241.

ENVR 5241. Lab for ENVR 5240. 1 Hour.

Accompanies ENVR 5240. Lab work uses geologic sections, suites of sedimentary rocks and thin sections, and drill cores and bore hole logs to interpret and analyze the geologic history and environmental and economic potential of sedimentary basins. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5240.

ENVR 5242. Ancient Marine Life. 4 Hours.

Begins with a survey of major events, processes, and important invertebrate phyla preserved in the fossil record. This knowledge of paleontology is then utilized to evaluate evolutionary principles and the nature of function and adaptation in the history of life. Organization of populations into paleocommunities and their relationships to changes in environments through time permit the assessment and evaluation of paleoecology in Earth history. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5243.

ENVR 5243. Lab for ENVR 5242. 1 Hour.

Accompanies ENVR 5242. Introduces invertebrate fossil morphology by study of fossil specimens of all major groups. Principles of paleoecology and evolutionary theory are illustrated by analysis of suites of fossil specimens. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5242.

ENVR 5244. Sedimentation. 4 Hours.

Describes the physical processes of sedimentation and their role in the interpretation of sedimentary environments. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5245.

ENVR 5245. Lab for ENVR 5244. 1 Hour.

Accompanies ENVR 5244. Concentrates on the interpretation and description of the physical properties of sediments and sedimentary environments. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5244.

ENVR 5248. Marine Geology. 4 Hours.

Compares the balance between major sedimentary and tectonic forces in ocean basins and margins to the resulting ocean form. Topics include origin of continental margins, shelf sedimentation and transport, deep-sea processes, and sediments. Evaluates resource development of OCS oil, sand and gravel, and manganese nodules. Prereq. Junior, senior, or graduate standing.

ENVR 5250. Geology and Land-Use Planning. 4 Hours.

Studies the causes and solutions of geologic environmental problems related to land use. Emphasizes geologic-based land-use planning solutions to problems related to landslides, ground subsidence, coastal erosion, stream erosion, flooding, soil erosion, and groundwater pollution. Assignments are based on actual examples requiring application of concepts covered in the course. Prereq. (a) ENVR 1200 (may be taken concurrently) or graduate standing and (b) ENGW 1111, ENGW 1102, ENGL 1111, ENGL 1102, or graduate standing; restricted to students with junior, senior, or graduate standing.

ENVR 5260. Geographical Information Systems. 4 Hours.

Examines geographical information systems (GIS), a way to input, store, analyze, and display spatial data (data with a geographic location). Introduces the major components and applications of this exciting new tool. Consists of two lectures and one laboratory period a week. Laboratory exercises introduce methods of data analysis as well as practical issues of how to manipulate various GIS software packages.

ENVR 5262. GIS Workshop. 2 Hours.

Studies the basic techniques of reflection and refraction seismology, gravity, aeromagnetic and heat-flow processes, and the information they provide on the structure, composition, and dynamics of the earth’s interior. Prereq. Junior, senior, or graduate standing.

ENVR 5270. Glacial and Quaternary History. 4 Hours.

Examines the environmental conditions conducive to forming glaciers, the processes of ice movement, glacial erosion, modes of deposition, and the resulting landforms created under and around glaciers. Introduces the natural climate change of the ice age cycles and the major events of the Quaternary period. Prereq. (a) ENVR 1112, ENVR 1200, or ENVR 5190 and (b) junior, senior, or graduate standing. Coreq. ENVR 5271.

ENVR 5271. Lab for ENVR 5270. 1 Hour.

Accompanies ENVR 5270. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing. Coreq. ENVR 5270.

ENVR 5290. Engineering Geology. 4 Hours.

Explores engineering geology, the interdisciplinary study of how geology is applied to engineering projects. Covers the application of geologic thought and geophysical methods to the site selection and planning of human-constructed features, such as foundations, landfills, highways, dams, tunnels, power plants, and mines. An individual research project augments class activities. Prereq. Junior, senior, or graduate standing.

ENVR 5300. Graduate Research. 4 Hours.

Offers an individual research project under the direction of a faculty member. Prereq. Junior, senior, or graduate standing.

ENVR 5400. Marine Science Policy and Ethics. 3 Hours.

Offers ethics training for a critical review of marine policies in the following topical areas: marine environmental ethics (conservation and preservation), conflicts of interest/research integrity, human subjects/mammal protections, ethical challenges in marine science modeling, ethics of fishing governance (marine conservation and regulations), sustainability models for marine sciences, data management, and new models of comanagement and community engagement with marine research. Reviews critical environmental policies affecting marine resources (NEPA, CERCLA, RCRA, Endangered Species, Marine Mammal Protection, and Oil Pollution acts, Magnuson-Stevens Act, etc.). Critically evaluates case studies and ethical review of coastal management for sustainability and pollution control, marine fisheries, and energy development. Prereq. Junior, senior, or graduate standing; restricted to students in the College of Science.

ENVR 5976. Directed Study. 1-4 Hours.

Offers independent study of a specific topic not normally contained in the regular course offerings but within the area of competence of a faculty member. Prereq. Junior, senior, or graduate standing.

ENVR 5978. Independent Study. 1-4 Hours.

Offers independent work under the direction of members of the department on a chosen topic. Course content depends on instructor. Prereq. Junior, senior, or graduate standing.

ENVR 5984. Research. 1-4 Hours.

Offers an opportunity to conduct research under faculty supervision. Prereq. Junior, senior, or graduate standing.

ENVR 6220. Applied Hydrology. 4 Hours.

Covers the origin, distribution, and flow of groundwater in permeable sediments and bedrock; hydrological and geological characteristics of aquifers; regional flow systems that emphasize rock structure, stratigraphy, and other aspects of the geological environment; principles of hydrogeologic mapping and analysis; and an introduction to well testing and well hydraulics. An individual research project augments class activities. Coreq. ENVR 6221.

ENVR 6221. Lab for ENVR 6220. 1 Hour.

Accompanies ENVR 6220. Covers topics from the course through various experiments. Coreq. ENVR 6220.

ENVR 6255. Introduction to Remote Sensing. 4 Hours.

Explores the fundamental concepts of remote sensing of the environment. Topics include digital imagery from spacecraft, conventional and high-altitude aerial photography, orthophotography production, and surface modeling systems. Offers hands-on experience with basic functions of industry-standard image-processing software.

ENVR 6962. Elective. 1-4 Hours.

Offers elective credit for courses taken at other academic institutions.

ENVR 6964. Co-op Work Experience. 0 Hours.

Provides eligible students with an opportunity for work experience.

ENVR 6966. Practicum. 1-4 Hours.

Provides eligible students with an opportunity for practical experience.

ENVR 6976. Directed Study. 1-4 Hours.

Offers independent study of a specific topic not normally contained in the regular course offerings but within the area of competence of a faculty member.

Ecology, Evolution, and Marine Biology Courses

EEMB 5130. Ecological Dynamics. 4 Hours.

Offers a comprehensive overview of mathematical and computational concepts needed to construct (meta)population, (meta)community, and (meta)ecosystem models. Focuses on how to mathematically derive and model processes (growth, trophic and nontrophic species interactions, dispersal, and environmental variability) to understand patterns of population abundance and species diversity. Emphasizes the mathematical tools required to analyze the dynamical behavior of ecological models (stability, invasion, graphical, and numerical analyses) and validate model predictions using empirical data (via maximum likelihood and optimization methods). Prereq. (a) MATH 1241, MATH 1251, MATH 1341 (any of which may be taken concurrently,) or graduate standing and (b) ENGW 1111, ENGW 1102, ENGL 1111, ENGL 1102, or graduate standing; restricted to juniors, seniors, and graduate students; sophomores admitted by permission of instructor. Coreq. EEMB 5131.

EEMB 5131. Lab for EEMB 5130. 1 Hour.

Accompanies EEMB 5130. Offers supervised lab sessions designed to show how the topics covered in the lectures can be addressed in industry-standard programming environments. Coreq. EEMB 5130.

EEMB 5303. Marine Biology Careers Seminar. 1 Hour.

Covers the information and tools needed to begin pursuing career opportunities in marine biology. Encourages students to explore a variety of career paths, construct résumés, contact potential employers for their internship and permanent positions. Presents invited speakers from state and federal agencies, and from private consulting firms, to talk about their work and career track. Prereq. Junior, senior, or graduate standing.

EEMB 5504. Biology of Corals. 3 Hours.

Focuses on the biology of Scleractinian reef-building corals and associated anthozoans found in coral reef ecosystems. Topics include systematics, anatomy, physiology, and population biology of corals, with an emphasis on the latest techniques employed by coral molecular biologists and physiologists. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5506. Biology and Ecology of Fishes. 3 Hours.

Presents an examination of the systematics, functional morphology, and behavioral, larval, and community ecology of reef fishes through lectures. Field and laboratory experiments focus on morphology, behavior, and community ecology of reef fishes. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5508. Marine Birds and Mammals. 2 Hours.

Studies principles of classification, anatomy, physiology, behavior, and evolution of seabirds and marine mammals. Also addresses conservation and protection of animals and essential habitat. Includes field trips to observe local species. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5509.

EEMB 5509. Lab for EEMB 5508. 1 Hour.

Accompanies EEMB 5508. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing. Coreq. EEMB 5508.

EEMB 5511. Adaptations of Aquatic Organisms. 3 Hours.

Explores the adaptive responses of marine organisms to variations in environmental factors. Focuses on physiological responses to a variety of natural and anthropogenic conditions. The laboratory component includes a combination of field and laboratory experiments. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5512. Tropical Terrestrial Ecology. 1 Hour.

Studies the animals, plants, and ecosystems of the new world tropics, with the community structure and diversity of terrestrial Jamaican habitats as an example. Includes field trips to lowland forests, carbonate caves, and the Blue Mountain mist-montane forest. The issue of land use and development vs. conservation is a recurring theme. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5514. Marine Ecology. 4 Hours.

Examines processes and interactions in ocean ecosystems. Topics include an introduction to major ocean ecosystems; the biotic and abiotic factors influencing the distributions, abundances, and interactions of marine organisms; and the transformation and flux of energy and matter in marine systems. Particularly emphasizes local coastal habitats, which are used to demonstrate quantitative field research methods. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5515.

EEMB 5515. Lab for EEMB 5514. 1 Hour.

Accompanies EEMB 5514. Covers topics from the course through various experiments. Coreq. EEMB 5514.

EEMB 5516. Oceanography. 4 Hours.

Offers an integrated overview of physical, chemical, biological, and geological processes operating in the world ocean. Seemingly unrelated topics like plate tectonics, oscillating currents and waves in the atmosphere, the activities of microbes and phytoplankton, and land-use practices in the middle of the continent have global reach and interact with each other in surprising yet understandable ways. Examines how new technologies have allowed stunning insights into global weather and climate, the deep sea, biodiversity, and how the biogeochemistry of the oceans can be measured and understood. Presents data use and analysis and formal reasoning used in marine science. Views the ocean as a “system of systems” where integration of experience from disparate disciplines is key. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5517.

EEMB 5517. Lab for EEMB 5516. 1 Hour.

Accompanies EEMB 5516. Offers experiential field and laboratory exercises in oceanography. The New England rocky intertidal, subtidal, wetlands, barrier islands, and dunes provide opportunities for field exercises in marine geology, physical oceanography, and marine ecology. Investigates processes affecting changes in the global ocean, such as ocean acidification; temperature stress in organisms; hydrodynamic drag and lift; suspension feeding; and the ecophysiology of reef corals, boreal invertebrates, and macroalgae. Coreq. EEMB 5516.

EEMB 5518. Ocean and Coastal Processes. 2 Hours.

Examines the coupling between physical and biological processes on coral reefs and adjacent habitats. Focuses on biophysical, oceanographic, and benthic-pelagic processes acting in coral reef and associated nearshore ecosystems. Specific topics include oceanographic forcing mechanisms, organismal biomechanics, hydrodynamics, and nutrient dynamics. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5520. Coral Reef Ecology. 2 Hours.

Examines the ecology and paleoecology of coral reefs. This course highlights the ecological importance of coral reefs and associated nearshore communities, ecosystem function, changes in reef biotas through geologic time, and the causes and consequences of reef degradation worldwide. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5522. Experimental Design Marine Ecology. 4 Hours.

Includes introduction to and application of observational methods in three local marine habitats, experimental design, statistical analysis, R statistical computing and graphics software, and principles of marine ecology. Combines lecture, hand-on research experience, and computer laboratory and includes reading and analyzing the scientific literature and developing research projects. At the end of the semester, students are expected to demonstrate an integrative mastery of course topics by writing a scientific manuscript about a class experiment. Seeks to prepare students for practicing ecology in new environments and to provide students with the foundational knowledge necessary for pursuing more complex concepts in experimental design, statistical analysis, and marine ecology. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5523.

EEMB 5523. Lab for EEMB 5522. 1 Hour.

Accompanies EEMB 5522. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing. Coreq. EEMB 5522.

EEMB 5524. Molecular Marine Biology. 3 Hours.

Uses molecular approaches (electrophoresis and DNA) to determine genetic relationships at the population and species level for the study of ecological and evolutionary questions. Techniques learned are applied to research projects. Prereq. (a) BIOL 2301 and junior or senior standing or (b) graduate standing.

EEMB 5526. Marine Microbial Ecology. 2 Hours.

Examines the diversity of marine microorganisms and recent advances in the area of microbial ecology. Emphasizes the structure and function of microbial food webs in marine communities. Prereq. Junior, senior, or graduate standing. Coreq. EEMB 5527.

EEMB 5527. Lab for EEMB 5526. 1 Hour.

Accompanies EEMB 5526. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing. Coreq. EEMB 5526.

EEMB 5528. Marine Conservation Biology. 3 Hours.

Examines several critical issues facing marine ecosystems, including invasive species, marine pollution and eutrophication, fisheries impacts, physical alteration of habitats, and global climate change. Offers students an opportunity to spend field time surveying intertidal and subtidal habitats within the San Juan Islands and Friday Harbor Marine Reserve and to conduct independent research projects. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5530. Molecular Ecology and Evolution. 4 Hours.

Exposes students to the molecular techniques and analyses used to examine the genetic relationships among individuals, populations, and species. Prereq. (a) BIOL 2301 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5531.

EEMB 5531. Lab for EEMB 5530. 1 Hour.

Accompanies EEMB 5530. Covers topics from the course through various experiments. Coreq. EEMB 5530.

EEMB 5532. Physiological and Molecular Marine Ecology. 3 Hours.

Explores the physiological responses of marine organisms to variations in environmental factors. Uses complementary techniques, including molecular and physiological approaches, to determine genetic relationships at the species and population level and elucidate the mechanistic basis of organismic responses to environmental conditions at the level of genes and gene products. Prereq. (a) Either BIOL 2311 or EEMB 2302 and junior or senior standing or (b) graduate standing.

EEMB 5534. Marine Invertebrate Zoology and Botany. 4 Hours.

Surveys the major groups of marine invertebrates, algae, and plants, in addition to their ecological roles and relationships. Offers students an opportunity to learn to identify these groups and understand the mechanisms they use to survive and adapt to changing oceans. Topics include ecological and evolutionary importance, ecosystem engineering, adaptive physiology, and climate change effects. Emphasizes interrelationships among major taxa. Hands-on learning includes field identification; visits to intertidal and subtidal marine environments; and specimen dissection, preparation, and cataloging. Offers students an opportunity to improve skills in reading and discussing scientific literature, experimental design, and scientific communication. Prereq. Junior, senior, or graduate standing; Three Seas students only; not open to students who have taken EEMB 5500 or EEMB 5502. Coreq. EEMB 5535.

EEMB 5535. Lab for EEMB 5534. 1 Hour.

Accompanies EEMB 5534. Covers topics from the course through various experiments. Coreq. EEMB 5534.

EEMB 5536. Ocean and Coastal Sustainability. 3 Hours.

Offers students advanced training in the expanding field of sustainability, with a combined focus on the practical aspects of systems management and the theoretical understanding of whole-systems design and resiliency. Seeks to train future leaders capable of creating innovative solutions to sustainability issues at local and global levels. Key interdisciplinary themes discussed include the social and political aspects of ocean and coastal sustainability (i.e., education and communication), sustainable development and ecosystem stability, the impacts of climate change on ocean and coastal resilience, and the economic and entrepreneurial possibilities in the field of sustainability. Prereq. Junior, senior, or graduate standing; Three Seas students only.

EEMB 5548. Sociobiology. 4 Hours.

Studies sociobiology, a field of biology that strives to understand the biological basis of social behavior in animals. Sociobiology is a multidisciplinary science, meshing together ethology (animal behavior), ecology, genetics, population biology, and comparative psychology, all within the conceptual framework of evolutionary theory. Why do animals live in societies? Why do animals cooperate? Why do they sometimes show extreme forms of altruism? What are the costs and benefits of group living? Reviews studies on nonhuman animals that demonstrate sociobiological principles by using a series of in-class activities, computer modeling assignments, interpretation of graphical and tabulated data, collection and statistical analyses of behavioral data, as well as the generation and presentation of research. Prereq. (a) BIOL 2301 and junior or senior standing or (b) graduate standing.

EEMB 5560. Entomology. 4 Hours.

Studies the biology of insects and related arthropods including their anatomy, morphology, physiology, development, taxonomy, ecology, behavior, and life histories. Includes field and laboratory study of insect biology. Prereq. (a) BIOL 2301 and junior or senior standing or (b) graduate standing. Coreq. EEMB 5561.

EEMB 5561. Lab for EEMB 5560. 1 Hour.

Accompanies EEMB 5560. Covers topics from the course through field and laboratory study, including insect collection. Prereq. Junior, senior, or graduate standing. Coreq. EEMB 5560.

EEMB 5562. Herpetology. 4 Hours.

Offers a survey of the amphibians and reptiles of the world, with emphasis on eastern North America. Topics include morphology, physiology, systematics, paleontology, ecology, zoogeography, and behavior. Includes field trips to observe the habitats and behavior of local herpetofauna. Laboratory emphasizes systematics and ecology. Prereq. (a) BIOL 1103, BIOL 1113, BIOL 2297, BIOL 2299, ENVR 2290, EEMB 2290, or graduate standing and (b) junior, senior, or graduate standing.

EEMB 5563. Lab for EEMB 5562. 1 Hour.

Accompanies EEMB 5562. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing.

EEMB 5564. Ornithology. 4 Hours.

Offers a survey of the birds of the world including morphology, physiology, systematics, behavior, ecology, zoogeography, and paleontology. Laboratory focuses on the identification and ecology of the avifauna of the Northeast, with field trips in eastern Massachusetts. Prereq. (a) BIOL 1103, BIOL 1113, BIOL 2297, BIOL 2299, ENVR 2290, EEMB 2290, or graduate standing and (b) junior, senior, or graduate standing.

EEMB 5565. Lab for EEMB 5564. 1 Hour.

Accompanies EEMB 5564. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing.

EEMB 5566. Mammalogy. 4 Hours.

Provides a survey of the mammals of the world including morphology, physiology, systematics, behavior, ecology, zoogeography, and paleontology. Laboratory focuses on the identification of the mammals of eastern North America and techniques used to study them. Includes a three-day field trip to observe mammals and employ techniques. Prereq. (a) BIOL 1103, BIOL 1113, BIOL 2297, BIOL 2299, ENVR 2290, EEMB 2290, or graduate standing and (b) junior, senior, or graduate standing.

EEMB 5567. Lab for EEMB 5566. 1 Hour.

Accompanies EEMB 5566. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing.

EEMB 5568. Wildlife Biology. 4 Hours.

Presents concepts and techniques utilized in the conservation and study of wild animals including the sociological aspects of management. Topics include habitat management, nonnative species, zoonoses, endangered species, legislation, and financing. Includes extended field trips to observe various ecosystems and wildlife. Prereq. (a) BIOL 1103, BIOL 1113, BIOL 2297, BIOL 2299, ENVR 2290, EEMB 2290, or graduate standing and (b) junior, senior, or graduate standing.

EEMB 5569. Lab for EEMB 5568. 1 Hour.

Accompanies EEMB 5568. Covers topics from the course through various experiments. Prereq. Junior, senior, or graduate standing.

EEMB 5589. Diving Research Methods. 2 Hours.

Presents experimental design, sampling methodology, statistical analysis, techniques, and the use of underwater equipment to conduct subtidal research. Prereq. Junior, senior, or graduate standing.

EEMB 6402. Concepts and Trends in Evolution and Ecology. 4 Hours.

Presents key concepts and important recent advances in evolution and ecology, including interdisciplinary approaches to understanding the distributions, abundances, and diversity of species, organisms, and molecules. Topics include natural selection, adaptation, speciation, molecular evolution, global change, and perspectives on communities and ecosystems. Discusses and critiques current literature and methods. Prereq. Biology and marine biology students only.

EEMB 7100. Colloquium. 1 Hour.

Offers a seminar-style course that includes weekly lectures and presentations of selected topics. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 8507. Marine Biology Graduate Co-op Tutorial. 1 Hour.

Designed to complement learning during co-op. Offers students an opportunity to participate in activities to integrate academic learning and experiential learning including written reflections. Helps students share their experiences in the workplace through class discussions moderated by the instructor.

EEMB 8674. Marine Biology Research Project. 1 Hour.

Offers an opportunity to design and implement a scientifically rigorous independent research project that builds upon current knowledge from the primary literature, under the supervision of a faculty advisor from the program. Students conduct research at any of the program’s locations and are then required to analyze data using rigorous statistical methods, write a journal-style research paper, and present their results in a research seminar.

EEMB 8960. Exam Preparation—Doctoral. 0 Hours.

Offers students an opportunity to prepare for the PhD qualifying exam under faculty supervision. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 8982. Readings. 1-4 Hours.

Assigns students independent readings on selected topics in ecology, evolution, and marine biology. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 8984. Research. 1-4 Hours.

Offers students an opportunity to conduct research. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 8986. Research. 0 Hours.

Offers students an opportunity to conduct full-time research under faculty supervision. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 9000. PhD Candidacy Achieved. 0 Hours.

Indicates successful completion of the doctoral comprehensive exam. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 9990. Dissertation. 0 Hours.

Offers theoretical and experimental research for the PhD degree. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.

EEMB 9996. Dissertation Continuation. 0 Hours.

Offers dissertation supervision by members of the department. Prereq. Restricted to PhD students in ecology, evolution, and marine biology.